In Santa Cecilia, Mexico, Miguel dreams of becoming a musician, even though his family strictly forbids it. His great-great-grandmother Imelda was married to a man who left her and their daughter Coco to pursue a career in music, and when he never returned, Imelda discarded all forms of music from her family’s life before starting a shoemaking business. Miguel now lives with the elderly Coco and their family, including Miguel’s parents and grandmother, who are all shoemakers. Coco suffers from memory loss and has become largely non-verbal, but Miguel is very close to her.
Miguel secretly idolizes famed musician Ernesto de la Cruz and practices his guitar skills using Ernesto’s old films. On the Day of the Dead, Miguel accidentally damages the picture frame that holds a photo of Coco with her mother and her father (the latter’s face torn off the picture) on the family ofrenda, discovering that a hidden section of the photograph shows his great-great-grandfather holding Ernesto’s famous guitar. Miguel, concluding that Ernesto is his great-great-grandfather, ignores his family’s objections and decides to enter the local talent show.
Breaking into Ernesto’s mausoleum, Miguel takes his guitar to use in the show, but once he strums it, he becomes invisible to everyone in the village plaza. However, he can interact with his skeletal deceased relatives, who are visiting from the Land of the Dead for the holiday.
Taking him back with them, they learn that Imelda cannot visit, since Miguel accidentally removed her photo from the ofrenda. Miguel discovers that he is cursed for stealing from the dead, and must return to the Land of the Living before sunrise, or he will become one of the dead; to do so, he must receive a blessing from a member of his family. Imelda offers Miguel a blessing on the condition of ending his dream of becoming a musician, but Miguel refuses and resolves to seek Ernesto’s blessing instead.
He meets Héctor, who declares that he knows Ernesto, offering to help Miguel reach him in return for Miguel taking his photo back with him, so that he might visit his daughter before she forgets him, causing him to disappear completely. Héctor helps Miguel enter a talent competition to win entry to Ernesto’s mansion, but Miguel’s family tracks him down, forcing him to flee. Miguel sneaks into the mansion, where Ernesto welcomes him as his descendant, but Héctor confronts them, again imploring Miguel to take his photo to the Land of the Living.
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