Remy is a highly intelligent and idealistic young rat with unusually acute senses of taste and smell who dreams of becoming a chef like his idol, the late Auguste Gusteau. However, the rest of his rat colony, including his brother Emile and his father Django, are interested in food only for sustenance. One day, when the colony is forced to flee their home, Remy is separated from the clan and eventually finds himself above the kitchen of Gusteau’s Restaurant in Paris.
When Remy observes a young garbage boy named Alfredo Linguini attempt to fix a soup he has spilled, he recognizes that Linguini is unintentionally ruining it and fixes his mistakes. Linguini catches him in the act, but does not reveal him to Skinner, Gusteau’s former sous-chef and new owner of the restaurant. Skinner confronts Linguini for tampering with the soup, but when the soup is accidentally served and proves to be a success, Colette Tatou, the staff’s only female chef, convinces Skinner to retain Linguini.
Skinner spots Remy trying to escape, and orders Linguini to kill the rat, but Linguini decides to keep Remy instead. When Linguini is ordered to replicate the soup, he is helped by Remy. Skinner, having tried the soup, decides to retain Linguini. Back in Linguini’s apartment the two learn to communicate and devise a plan: Remy hides under Linguini’s toque at the restaurant and guides his movements like a marionette by pulling on his hair. Skinner orders Colette to teach Linguini to be a cook.
Remy witnesses Skinner’s discovery that Linguini is Gusteau’s illegitimate son and rightful owner of the restaurant, and gives the evidence to Linguini, who forces Skinner out. The restaurant thrives as Remy’s recipes become popular, though as Linguini and Colette develop a relationship, Remy begins to feel left out. He reunites with Emile and the clan, but is rejected by Django over his admiration for humans.
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