In the 29th century, rampant consumerism and environmental neglect have turned Earth into a garbage-strewn wasteland; humanity is nowhere to be found and has been evacuated by the megacorporation Buy n Large (BnL) on giant starliners seven centuries earlier. Of all the Waste Allocation Load-Lifter: Earth-Class robots left by BnL to clean up, only one named WALL-E remains operational.
One day, WALL-E’s routine of compressing trash and collecting interesting objects is broken by the arrival of an unmanned probe carrying an Extraterrestrial Vegetation Evaluator robot named EVE, sent to scan the planet for human-sustainable life. WALL-E is smitten by the sleek, otherworldly robot; and the two begin to connect until EVE goes into standby when WALL-E shows her his most recent find: a living seedling. The probe then collects EVE and the plant, and—with WALL-E clinging on—returns to its mothership, the starliner Axiom.
In the centuries since the Axiom left Earth, its passengers have degenerated into helpless corpulence due to laziness and microgravity, their every whim catered to by machinery; even the captain, one B. McCrea, is used to sitting back while his robotic lieutenant and steering wheel AUTO flies the ship. McCrea is taken aback by a positive probe response and learns that placing the plant in the ship’s Holo-Detector will trigger a hyperjump back to Earth so humanity can begin recolonization. Upon inspecting EVE’s storage compartment, however, the plant turns up missing, and EVE blames WALL-E for its disappearance.
With the plant missing, EVE is deemed faulty and taken to Diagnostics. Mistaking the process for torture, WALL-E shuts all of Diagnostics down, causing him and EVE to be designated as rogues. Frustrated, EVE tries to send WALL-E home on an escape pod, but they are interrupted when AUTO’s first mate GO-4 arrives and stows the stolen plant in a pod set to self-destruct. WALL-E saves the plant from being lost for good, and he and EVE reconcile and celebrate with a “dance in space” around the Axiom.
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